Family Orchidaceae Subfamily Orchidoideae Tribe Cranichideae Subtribe Goodyerinae

Family Orchidaceae Subfamily Orchidoideae Tribe Cranichideae Subtribe Goodyerinae

Goodyerinae Klotzsch (as ‘Goodyereae’) in Hayne, Getreue Darstell. Gew. (1846) 14, t. 24. Type genus: Goodyera R.Br.


  • Tribe Goodyerinae King & Pantl., Ann. Roy. Bot. Gard. Calcutta, 8 (1898) 273.
  • Subtribe Physurinae Lindl., Gen: Sp. Orch. Pl., 442 (1840), [as Physuridae]. Type genus: Physurus L.C.Rich. ex Lindl., nom. illeg.
  • Monandreae Neottiinae Physureae (Lindl.) Pfitzer, in Engler & Prantl, Die Natürl. Pflanzenf. II. 6. Orchidaceae (1889) n4. Type genus: Physurus L.C.Rich. ex Lindl., nom. illeg.
  • Subtribe Pachyplectroninae Schltr. Notizbl. Bot. Gart. Mus. Berlin-Dahlem 9 (1926) 570. Type genus: Pachyplectron Schltr.
  • Subtribe Cheirostylidinae Szlach., Folia Geobot. Phytotax. 26 (1991) 318. Type genus: Cheirosrylis Blume.

Perennial terrestrials, sometimes lithophytes and rarely epiphytic, herbaceous. Roots usually fleshy, hairy, from the nodes of the rhizome, rarely absent and replaced by rhizoids; velamen present or not. Rhizome creeping, short to elongate; internodes few to many, of equal length, rarely some abbreviated. Stem erect, terete, few- to many-leaved, rarely leafless. Leaves sessile, rarely reduced to sheaths and then without chlorophyll, or with a tubular leaf sheath and with a petiole-like base; leaf blade obliquely lanceolate to subcircular. lnflorescence terminal, erect, glabrous or with indument; peduncle usually with a few scale leaves; floral floral bracts lanceolate to ovate glabrous, glandular, or pubescent. Flowers resupinate or rarely not, glabrous or the sepals outside glandular or pubescent. Ovary cylindrical-fusiform, glabrous, glandular or pubescent. Sepals usually, rarely connate at the base. Median sepal concave, often forming a hood with the petals. Lateral sepals free or connate, often oblique at the base; lower half clasping the lip. Petals free or connate entire, sessile or with a claw, often hyaline, usually adnate to the dorsal sepal. Lip usually fleshy; attached to the base and usually also to the base of the column, entire, 2- or 3-lobed; hypochile concave, saccate or spurred, sometimes with a fleshy external flange; inside on both sides with or without 1 or few finger-shaped, stalked or sessile appendages, rarely all over the inside, sometimes with 2 keels; epichile sessile or clawed, the claw fringed or not, small to large, linear, ovate, moon-shaped or transversely 2-lobed; spur absent or present, entire or 2-lobed, tapering, cylindrical or club-shaped. Column short to long, sessile or with a stalk; anther usually erect, 2-loculate; pollinia 2, 2-partite or rarely entire, sectile, usually club-shaped, attenuated into short to elongate caudicles; clinandrium usually deeply concave; rostellum short to elongate, erect to porrect, often deltoid, slightly emarginate to deeply bifid, in some genera carrying a distinct, dorsal, stipes; stigma lobes free or connate under the base of the rostellum, sometimes subtended by 1 or 2 sterile appendages. Fruit usually cylindrical-fusiform, dehiscent.
(after Ormerod & Cribb 2003)

Subtribe Goodyerinae contains 34 genera. It is widely distributed in both the Old and New World, particularly in the tropics and subtropics: Europe, Asia, Australasia, Africa, Macronesia, Madagascar, and the Americas. The genera may be found as far north as Canada, Russia, Japan, and Korea, and as far south as Australia, New Zealand, Argentina, and South Africa.

  • Ormerod. P. & Cribb. P.J., in Pridgeon, A.M., P.J. Cribb, M.W. Chase & F.N. Rasmussen (edit.), Genera Orchidacearum 3, Orchidoideae (part 2) (2003) 63
  • Family Orchidaceae
  • Subfamily Orchidoideae
  • Tribe Cranichideae
  • Subtribe Goodyerinae

Subtribe Goodyerinae in New Guinea contains 16 genera:



1a. Flowers not-resupinate == 2
1b. Flowers resupinate == 5

2a. Stigma not lobed == 3
2b. Stigma with 2 lobes == 4

3a. Column in front without wings. Lip hypochile oblong-elliptic == Papuaea
3b. Column in front at the base with quadrate to rounded wings. Lip hypochile subquadrate == Macodes

4. Leaves with brightly coloured nerves == Anoectochilus
4b. Leaves without brightly coloured nerves == Hetaeria

5a. Stigma with 2 distinct lobes == 6
5b. Stigma entire == 11

6a. Lip with a cylindrical to fusiform spur == 7
6b. Lip without spur == 8

7. Column in front with wings. Leaves with brightly coloured nerves == Anoectochilus
7b. Column in front without wings. Leaves without brightly coloured nerves == Vrydagzynea

8a. Lip with a raised median keel == Rhomboda
8b. Lip without a raised median keel == 9

9a. Stigma lobes not stalked == 10
9b. Stigma lobes stalked == Myrmechis

10a. Stigma lobes without a projecting appendage == Zeuxine
10b. Stigma lobes with a projecting appendage == Cheirostylis

11a. Spur or hypochile of the lip not enclosed by the widely opened lateral sepals == 12
11b. Spur or hypochile of the lip enclosed by the base of the lateral sepals, rarely somewhat exposed == 15

12a. Spur inside with several randomly scattered appendages == Goodyera
12b. Spur inside with 2 appendages situated on the intraneural veins, or inside without appendages == 13

13a. Column without raised stigma lobes, semiterete to obliquely conical == Erythrodes
13b. Column with raised stigma lobes with marginate edges, appearing semi-malleolate to dolabriform in lateral view == 14

14a. Lateral sepals at the base saccate, enclosing the inflated bladders of the spur. == Cystorchis
14b. Lateral sepals at the base not saccate; spur ewithout inflated bladders== Eurycentrum

15a. Plants heteromycotrophic. Leaves reduced to scales == Cystorchis
15b. Plants autotrophic. Leaves normally developed == 16

16a. Leaves fleshy, elongate, ligulate. Lip apex ligulate; hypochile inside with a transverse row of calli at the base == Lepidogyne
16b. Leaves not fleshy, ovate to elliptic-lanceolate. Lip apex not ligulate; hypochile inside without a transverse row of calli == 17

17a. Lip with entire to toothed external flanges on the mesochile or hypochile == Kuhlhasseltia
17b. Lip without entire to toothed external flanges on the mesochile or hypochile == 34

18a. Lip inflated saccate == Hylophila
18b. Lip not inflated saccate == 19

19a. Lip hypochile median or below the middle somewhat constricted == Platylepis
19b. Lip hypochile not constricted == Goodyera

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