Species Information - Pachystoma pubescens

Pachystoma pubescens Blume, Bijdr. (1825) 376; Tab. & Pl. Jav. Orchid. (1825) t. 29

Type: Blume s.n. (Tjiradjas, Krawang Prov.) (holo L, possibly, the only Blume specimen in L is Blume 1529 annotated as 'Pachichilus javanicus' Blume)

Synonyms:

• Pachychilus pubescens (Blume) Blume, Mus. Bot. Lugd. Bat. 2 (1856) 173
• Apaturia chinensis Lindl., Gen. Sp. Orchid. Pl. (1831) 131
  - Type: Vachel s.n. (Chia, Macao)
• Apaturia montana Lindl., Gen. Sp. Orchid. Pl. (1831) 131
  - Type: Wallich 3738 (India) (K); Macrae s.n. (Ceylon, Mt. Nuera Ellia) (K)
• Pachystoma montanum (Lindl.) Rchb.f., Bonplandia 3 (1855) 251
• Pachystoma senile (Lindl.) Rchb.f., Bonplandia 3 (1858) 251
• Pachychilus senilis (Lindl.) Blume, Mus. Bot. Lugd. Bat. 2 (1856) 173
• Apaturia senilis Lindl., Gen. Sp. Orchid. Pl. (1831) 130
  - Type: Wallich (Bruce) 3739 (India, Syllet) (lecto K, chosen by Clements 1989; isolecto K-WALL); Hamilton s.n. (Uligapur and Kalegung) (syn K, K, K-WALL)
• Apaturia smithiana Lindl., Gen. Sp. Orchid. Pl. (1831) 131
  - Type: Wallich 3737 (Kudjura-ghaut, Oude) (holo K; iso K-WALL, L)
• Pachystoma smithianum (Lindl.) Rchb.f., Bonplandia 3 (1858) 251
• Pachystoma lindleyanum (Wight) Rchb.f., Bonplandia 3 (1855) 251
• Apaturia lindleyana Wight, Icon. Pl. Ind. Or. 5 (1851) 8, t. 1662
  - Type: Jerdon s.n. (India, Coorg) (holo K; icon. K)
• Pachystoma edgeworthii Rchb.f. in Walp., Ann. Bot. Syst. 3 (1853) 464
  - Type: Edgeworth 25 (India NW, Dyanhar, Khadia) (holo K; illustr. K)
• Neottia pantanis Korth. ex Blume, Mus. Bot. Lugd. Bat. 2 (1856) 173 (in syn.)
• Pachystoma pantanum (Blume) Miq., Fl. Ned. Indiñ3 (1859) 675
• Pachychilus pantanus Blume, Mus. Bot. Lugd. Bat. 2 (1856) 173
  - Type: Korthals s.n. (Borneo, Padong antam) (holo L 2x; iso K)
• Eulophia holtzei F.Muell., Victorian Naturalist 6 (1889) 55
  - Type: Holtze s.n. (Australia Port Darwin) (holo K)
• Pachystoma gracile Schltr., Bot. Jahrb. Syst. 39 (1906) 66
  - Type: Schlechter 15752 (New Caledonia N, Ou Hinna) (holo B, lost)
• Pachystoma pubescens Blume var. gracile (Schltr.) N.Hallé, Fl. Nouv.-Caléd. Dép. 8 (1977) 205
• Pachystoma chinensis Hayata, J. Coll. Sci. Imp. Univ. Tokyo 30 (1911) 321 (non Rchb.f.)
  - Type: Kawakami & Mori 3735 (Taiwan, Nanto, Shashizan) (holo TI)
• Pachystoma affine Schltr., Repert. Spec. Nov. Regni Veg. Beih. 1 (1912) 413
  - Type: Schlechter 16573 (PNG, 'Kaiser Wilh. Land', Minjem R., Jawer) (holo B, lost)
• Pachystoma papuanum Schltr., Repert. Spec. Nov. Regni Veg. Beih. 1 (1912) 412
  - Type: Schlechter 18472 (PNG, foor Bismarck Ra.) (holo B, lost)
• Pachystoma brevilabium Schltr., Repert. Spec. Nov. Regni Veg. 17 (1921) 69
  - Type: Esquirol s.n. (VI-1912; Lo-kouey, Kweitschou) (holo B, lost)
• Pachystoma chinense Rchb.f. var. formosanum (Schltr.) S.S.Ying, Col. Illust. Indig. Orchid. Taiwan 1, 2 (1977) 488
• Pachystoma formosanum Schltr., Repert. Spec. Nov. Regni Veg. Beih. 4 (1919) 245 (as nom. nov.)
  - Type: ?
• Pachystoma hirsuta (J.Joseph & Vajr.) C.S.Kumar & Manilal, Kew Bull. 42 (1987) 942
• Eulophia hirsuta J.Joseph & Vajr., Bull. Bot. Surv. India 17 (1975 publ. 1978) 192, fig. 1-8a
  - Type: Vajravelu 46201 (holo CNH; iso Rapinat herb; Kanjilal herb.)

Rhizome horizontal, fleshy, c. 6 cm long. Leaves absent at flowering time, 1 or 2, narrowly linear, c. 30 by 0.7 cm, acuminate. Inflorescence erect, elongated, laxly c. 6-15-flowered, with the flowers more or less turned to one side; peduncle 42-56 cm long; peduncle-scales tubular, acute, becoming smaller towards the apex, to c. 7.5 cm long; rachis very sparsely pubescent, 5-11 cm long. Floral bracts linear-lanceolate, gradually acuminate, at the base dilated and slightly puberulous, somewhat transparent, membranous, 7-nerved, to 3 cm long, upper ones smaller. Sepals outside puberulous. Median sepal oblong, obtuse, concave, 1.25 by 0.46 cm. Lateral sepals obliquely oblong, shortly acuminate, acute, especially near the base concave, 5-nerved, 1.35 by 0.43 cm. Petals linear-lanceolate, towards the apex somewhat dilated, obtuse, apiculate, 3-nerved, outside especially near the base as well as inside sparsely puberulous, very sparsely ciliolate, 1.3 by 0.23 cm. Lip at the base together with the column-foot forming a short sac enveloping the base of the lateral sepals, clawed, 3-lobed, when flattened 1.2 cm by 0.9-1.1 cm, at the base with a glabrous concavity delimited by a transverse thickening, in the basal part inside and outside puberulous and with two basally puberulous keels, in the middle with three keels, at the base of the mid-lobe with two short additional keels, all keels warty, the median keel longest; lateral lobes erect, surpassing the column, broadly obliquely subovate, broadly obtuse, inside very sparsely puberulous; mid-lobe square-obcordate or almost quadrangular, 0.4 cm long and wide, emarginate, sometimes with a triangular apicule in the sinus, crenate, inside with a few scattered hairs. Column very slender, 0.75 cm long, making an obtuse angle with the ovary, curved, puberulous, at the apex subglobose-swollen, 3-lobulate, median lobule (filament) truncate, almost bilobulate, lateral lobules (auricles) larger, broad, rounded, outside convex; clinandrium broad, deeply concave; stigma large, transverse, deeply concave, lower margins thin; column-foot very short, concave, at the base with a densely puberulous thickening, 0.15 cm long. Anther subreniform, slightly puberulous; pollinia 8, laterally flattened, pear-shaped. Pedicel and ovary velvety hairy, ovary 0.6-0.7 cm long, pedicel much more slender, 0.2-0.3 cm long. (After Smith, 1909)

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